As I have alluded more than once in this chapter to the occurrence of the remains of the hippopotamus in places where there are now no rivers, not even a rill of water, and as other bones of the same genus have been met with in the lower-level gravels of the Somme where large blocks of sandstone seem to imply that ice once played a part in their transportation, it may be well to consider, before proceeding farther, what geographical and climatal conditions are indicated by the presence of these fossil pachyderms.
It is now very generally conceded that the mammoth and tichorhine rhinoceros were fitted to inhabit northern regions, and it is therefore natural to begin by asking whether the extinct hippopotamus may not in like manner have flourished in a cold climate. In answer to this inquiry, it has been remarked that the living hippopotami, anatomically speaking so closely allied to the extinct species, are so aquatic and fluviatile in their habits as to make it difficult to conceive that their congeners could have thriven all the year round in regions where, during winter, the rivers were frozen over for months. Moreover, I have been unable to learn that, in any instance, bones of the hippopotamus have been found in the drift of northern Germany associated with the remains of the mammoth, tichorhine rhinoceros, musk-ox, reindeer, lemming, and other arctic quadrupeds before alluded to; yet, though not proved to have ever made a part of such a fauna, the presence of the fossil hippopotamus north of the fiftieth parallel of latitude naturally tempts us to speculate on the migratory powers and instincts of some of the extinct species of the genus. They may have resembled, in this respect, the living musk-ox, herds of which pass for hundreds of miles over the ice to the rich pastures of Melville Island, and then return again to southern latitudes before the ice breaks up.
We are indebted to Sir Andrew Smith,* an experienced zoologist, for having given us an account of the migratory habits of the living hippopotamus of Southern Africa (H. amphibius, Linn.).
(* "Illustrations of the Zoology of South Africa": article
"Hippopotamus.")
He states that, when the Dutch first colonised the Cape of Good Hope, this animal abounded in all the great rivers, as far south as the land extends; whereas, in 1849, they had all disappeared, scarcely one remaining even within a moderate distance of the colony. He also tells us that this species evinces great sagacity in changing its quarters whenever danger threatens, quitting every district invaded by settlers bearing fire-arms. Bulky as they are, they can travel speedily for miles over land from one pool of a dried-up river to another; but it is by water that their powers of locomotion are surpassingly great, not only in rivers, but in the sea, for they are far from confining themselves to fresh water. Indeed, Sir A. Smith finds it "difficult to decide whether, during the daytime and when not feeding, they prefer the pools of rivers or the waters of the ocean for their abode." In districts where they have been disturbed by Man, they feed almost entirely in the night, chiefly on certain kinds of grass, but also on brushwood. Sir A. Smith relates that, in an expedition which he made north of Port Natal, he found them swarming in all the rivers about the tropic of Capricorn. Here they were often seen to have left their footprints on the sands, entering or coming out of the salt water; and on one occasion Smith's party tried in vain to intercept a female with her young as she was making her way to the sea. Another female, which they had wounded on her precipitate retreat to the sea, was afterwards shot in that element.
The geologist, therefore, may freely speculate on the time when herds of hippopotami issued from North African rivers, such as the Nile, and swam northwards in summer along the coasts of the Mediterranean, or even occasionally visited islands near the shore. Here and there they may have landed to graze or browse, tarrying awhile and afterwards continuing their course northwards. Others may have swum in a few summer days from rivers in the south of Spain or France to the Somme, Thames, or Severn, making timely retreat to the south before the snow and ice set in.